Publications

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Christiansen, G. B., Petersen, L. V., Chatwin, H., Yilmaz, Z., Schendel, D., Bulik, C. M., Grove, J., Brikell, I., Semark, B. D., Holde, K., Abdulkadir, M., Hübel, C., Albiñana, C., Vilhjálmsson, B. J., Børglum, A. D., Demontis, D., Mortensen, P. B. & Larsen, J. T. (2025). The role of co-occurring conditions and genetics in the associations of eating disorders with attention-deficit/hyperactivity disorder and autism spectrum disorder. Molecular Psychiatry, 30(5), 2127-2136. Article 132. https://doi.org/10.1038/s41380-024-02825-w

Pathi, A., Wright, M., Smed, M. K., Nelson, J. L., Olsen, J., Hetland, M. L., Zoffmann, V. & Jawaheer, D. (2021). The rheumatoid arthritis gene expression signature among women who improve or worsen during pregnancy: A pilot study. Journal of Rheumatology, 48(7), 985-991. https://doi.org/10.3899/jrheum.201128

Ahrenfeldt, J., Christensen, D. S., Østergaard, A. B., Kisistók, J., Sokač, M. & Birkbak, N. J. (2023). The ratio of adaptive to innate immune cells differs between genders and associates with improved prognosis and response to immunotherapy. PLoS One, 18(2 ), Article e0281375. https://doi.org/10.1371/journal.pone.0281375

Christensen, D. S. & Birkbak, N. J. (2023). Therapy drives genomic evolution in metastatic cancer. OncoTarget, 14(1), 216-218. https://doi.org/10.18632/oncotarget.28379

Schoustra, S. E., Bataillon, T., Gifford, D. R. & Kassen, R. (2009). The properties of adaptive walks in evolving populations of fungus. PLoS Biology, 7(11), e1000250. https://doi.org/10.1371/journal.pbio.1000250

Wiuf, C., Zhao, K., Innan, H. & Nordborg, M. (2004). The probability and chromosomal extent of trans-specific polymorphisms. Genetics, 168, 2363-2372.

Castagnini, A., Foldager, L., Caffo, E. & Berrios, G. E. (2022). The predictive validity and outcome of ICD-10 and DSM-5 short-lived psychotic disorders: a review and meta-analysis. European Archives of Psychiatry and Clinical Neuroscience, 272(7), 1157-1168. https://doi.org/10.1007/s00406-021-01356-7

Christiansen, F. (2008). Theories of Population Variation in Genes and Genomes. Princeton University Press.

Kong, A., Thorleifsson, G., Frigge, M. L., Vilhjalmsson, B. J., Young, A. I., Thorgeirsson, T. E., Benonisdottir, S., Oddsson, A., Halldorsson, B. V., Masson, G., Gudbjartsson, D. F., Helgason, A., Bjornsdottir, G., Thorsteinsdottir, U. & Stefansson, K. (2018). The nature of nurture: Effects of parental genotypes. Science, 359(6374), 424-428. https://doi.org/10.1126/science.aan6877

Skov, L., Coll Macià, M., Sveinbjörnsson, G., Mafessoni, F., Lucotte, E. A., Einarsdóttir, M. S., Jonsson, H., Halldorsson, B., Gudbjartsson, D. F., Helgason, A., Schierup, M. H. & Stefansson, K. (2020). The nature of Neanderthal introgression revealed by 27,566 Icelandic genomes. Nature, 582(7810), 78-83. https://doi.org/10.1038/s41586-020-2225-9

Bergeron, L. A., Besenbacher, S., Turner, T. N., Versoza, C. J., Wang, R. J., Price, A. L., Armstrong, E., Riera, M., Carlson, J., Chen, H. Y., Hahn, M. W., Harris, K., Kleppe, A. S., López-Nandam, E. H., Moorjani, P., Pfeifer, S. P., Tiley, G. P., Yoder, A. D., Zhang, G. & Schierup, M. H. (2022). The mutationathon highlights the importance of reaching standardization in estimates of pedigree-based germline mutation rates. eLife, 11, Article e73577. https://doi.org/10.7554/eLife.73577

Hobolth, A. & Siren, J. (2016). The multivariate Wright–Fisher process with mutation: Moment-based analysis and inference using a hierarchical Beta model. Theoretical Population Biology, 108, 36-50. https://doi.org/10.1016/j.tpb.2015.11.001

Christiansen, F. (1989). The multiple locus symmetric fertility model. Theoretical Population Biology, 35, 337-362.

Murat, F., Mbengue, N., Winge, S. B., Trefzer, T., Leushkin, E., Sepp, M., Cardoso-Moreira, M., Schmidt, J., Schneider, C., Mößinger, K., Brüning, T., Lamanna, F., Riera Belles, M., Conrad, C., Kondova, I., Bontrop, R., Behr, R., Khaitovich, P., Pääbo, S. ... Kaessmann, H. (2023). The molecular evolution of spermatogenesis across mammals. Nature, 613(7943), 308-316. https://doi.org/10.1038/s41586-022-05547-7

Stukenbrock, E. H., Bataillon, T., Dutheil, J. Y., Hansen, T. T., Li, R., Zala, M., McDonald, B. A., Wang, J. & Schierup, M. H. (2011). The making of a new pathogen: Insights from comparative population genomics of the domesticated wheat pathogen Mycosphaerella graminicola and its wild sister species. Genome Research, 21, 2157-2166. https://doi.org/10.1101/gr.118851.110

Malolepszy, A., Mun, T., Sandal, N., Gupta, V., Dubin, M., Urbanski, D. F., Shah, N., Bachmann, A., Fukai, E., Hirakawa, H., Tabata, S., Nadzieja, M., Markmann, K., Su, J., Umehara, Y., Soyano, T., Miyahara, A., Sato, S., Hayashi, M. ... Andersen, S. U. (2016). The LORE1 insertion mutant resource. The Plant Journal, 88(2), 306-317. https://doi.org/10.1111/tpj.13243

Montes, M., Nielsen, M. M., Maglieri, G., Jacobsen, A., Hojfeldt, J., Agrawal-Singh, S., Hansen, K., Helin, K., de Werken, H. J. G. V., Pedersen, J. S. & Lund, A. H. (2015). The lncRNA MIR31HG regulates p16(^INK4A) expression to modulate senescence. Nature Communications, 6, Article 6967. https://doi.org/10.1038/ncomms7967

Schierup, M. H. (2020). The last pieces of a puzzling early meeting. Science (New York, N.Y.), 369(6511), 1565-1566. https://doi.org/10.1126/science.abe2766

Gao, H., Hamp, T., Ede, J., Schraiber, J. G., McRae, J., Singer-Berk, M., Yang, Y., Dietrich, A. S. D., Fiziev, P. P., Kuderna, L. F. K., Sundaram, L., Wu, Y., Adhikari, A., Field, Y., Chen, C., Batzoglou, S., Aguet, F., Lemire, G., Reimers, R. ... Farh, K. K. H. (2023). The landscape of tolerated genetic variation in humans and primates. Science (New York, N.Y.), 380(6648), Article abn8197. https://doi.org/10.1126/science.abn8197

Juul, R. I., Nielsen, M. M., Juul, M., Feuerbach, L. & Pedersen, J. S. (2021). The landscape and driver potential of site-specific hotspots across cancer genomes. npj Genomic Medicine, 6(1), Article 33. https://doi.org/10.1038/s41525-021-00197-6

Mailund, T. (2019). The Joys of Hashing: Hash Table Programming with C. Apress. https://doi.org/10.1007/978-1-4842-4066-3

Pedersen, C. B., Bybjerg-Grauholm, J., Pedersen, M. G., Grove, J., Agerbo, E., Bækvad-Hansen, M., Poulsen, J. B., Hansen, C. S., McGrath, J. J., Als, T. D., Goldstein, J. I., Neale, B. M., Daly, M. J., Hougaard, D. M., Mors, O., Nordentoft, M., Børglum, A. D., Werge, T. & Mortensen, P. B. (2018). The iPSYCH2012 case-cohort sample: new directions for unravelling genetic and environmental architectures of severe mental disorders. Molecular Psychiatry, 23(1), 6–14. https://doi.org/10.1038/mp.2017.196

Vodolazs'ka, D., Feyera, T., Foldager, L. & Lauridsen, C. (2023). The influence of supplementary feeding during suckling and later weaning on growth performance, incidences of post-weaning diarrhoea and immunity of piglets. Livestock Science, 276, Article 105321. https://doi.org/10.1016/j.livsci.2023.105321

Waltoft, B. L., Pedersen, C. B., Nyegaard, M. & Hobolth, A. (2015). The importance of distinguishing between the odds ratio and the incidence rate ratio in GWAS. BMC Medical Genetics, 16(1), 71. https://doi.org/10.1186/s12881-015-0210-1

Coutant, M. P., Malmkvist, J., Foldager, L. & Herskin, M. S. (2023). The importance of carefully selecting vocalization parameters to quantify pain and stress in piglets. In Book of Abstracts - ISAE 2023: 56th Congress of the International Society for Applied Ethology ISAE 2023 (pp. 68-68).

Rafnar, T., Thorlacius, S., Steingrimsson, E., Schierup, M. H., Madsen, J. N., Calian, V., Eldon, B. J., Jonsson, T., Hein, J. & Thorgeirsson, S. S. (2004). The Icelandic Cancer Project--a population-wide approach to studying cancer. Nat Rev Cancer Jun, 4(6), 488-492.

Bergeron, L. A., Besenbacher, S., Bakker, J., Zheng, J., Li, P., Pacheco, G., Sinding, M. H. S., Kamilari, M., Gilbert, M. T. P., Schierup, M. H. & Zhang, G. (2021). The germline mutational process in rhesus macaque and its implications for phylogenetic dating. GigaScience, 10(5), Article giab029. https://doi.org/10.1093/gigascience/giab029

Ma, J., Bechsgaard, J. S., Lauridsen, A. A., Villesen, P., Bilde, T. & Schierup, M. H. (2025). The genomic consequences and persistence of sociality in spiders. Genome Research, 35(3), 499-511. https://doi.org/10.1101/gr.279503.124

Wang, X., Nursyifa, C., Aninta, S. G., Garcia-Erill, G., Bertola, L. D., Khan, A., Kuja, J., Hanghøj, K., Meisner, J., Bøggild, T., Bradshaw, C. J. A., Al-Chaer, A., Persada, A. P., Priyono, D. S., Tribudi, Y. A., Sudrajad, P., Gaina, C. D., Jiang, Y., Lenstra, J. A. ... Heller, R. (2025). The genetic diversity of Indonesian cattle has been shaped by multiple introductions and adaptive introgression. Nature Communications, 16(1), Article 8192. https://doi.org/10.1038/s41467-025-62692-z

Christiansen, F. (1990). The generalized multiplicative model for viability selection at multiple loci. Journal of Mathematical Biology, 29, 99-129.

Hobolth, A. & Wiuf, C. H. (2009). The genealogy, site frequency spectrum and ages of two nested mutant alleles. Theoretical Population Biology, 75(4), 260-265. https://doi.org/10.1016/j.tpb.2009.02.001

Brown, P. E., Christensen, O. F., Clough, H. E., Diggle, P. J., Hart, C. A., Hazel, S., Kemp, R., Leatherbarrow, A. J. H., Moore, A., Sutherst, J., Turner, J., Williams, N. J., Wright, E. J. & French, N. P. (2004). The frequency and spatial distribution of environmental campylobacter spp. Applied Environmental Microbiology, 70, 6501-6511.

Fragata, I., Matuszewski, S., Schmitz, M. A., Bataillon, T., Jensen, J. D. & Bank, C. (2018). The fitness landscape of the codon space across environments. Heredity, 121(5), 422-437. https://doi.org/10.1038/s41437-018-0125-7

Wigdor, E. M., Weiner, D. J., Grove, J., Fu, J. M., Thompson, W. K., Carey, C. E., Baya, N., van der Merwe, C., Walters, R. K., Satterstrom, F. K., Palmer, D. S., Rosengren, A., Bybjerg-Grauholm, J., Hougaard, D. M., Mortensen, P. B., Daly, M. J., Talkowski, M. E., Sanders, S. J., Bishop, S. L. ... iPSYCH Consortium (2022). The female protective effect against autism spectrum disorder. Cell Genomics, 2(6), Article 100134. https://doi.org/10.1016/j.xgen.2022.100134

Feldman, M., Christiansen, F. & Holsinger, K. (1984). The evolution of self-pollination in plants. American Naturalist, 124, 446–453.

Morgan, M. T., Schoen, D. J. & Bataillon, T. (1997). The evolution of self-fertilization in perennials. American Naturalist, 150(5), 618-38. https://doi.org/10.1086/286085

Bach, L. A., Helvik, T. & Christiansen, F. B. (2006). The evolution of n-player cooperation—threshold games and ESS bifurcations. Journal of Theoretical Biology, 238, 426-434.

Frankell, A. M., Dietzen, M., Al Bakir, M., Lim, E. L., Karasaki, T., Ward, S., Veeriah, S., Colliver, E., Huebner, A., Bunkum, A., Hill, M. S., Grigoriadis, K., Moore, D. A., Black, J. R. M., Liu, W. K., Thol, K., Pich, O., Watkins, T. B. K., Naceur-Lombardelli, C. ... TRACERx Consortium (2023). The evolution of lung cancer and impact of subclonal selection in TRACERx. Nature, 616(7957), 525-533. https://doi.org/10.1038/s41586-023-05783-5

Hartfield, M., Bataillon, T. & Glemin, S. (2017). The Evolutionary Interplay between Adaptation and Self-Fertilization. Trends in Genetics, 33(6), 420-431. https://doi.org/10.1016/j.tig.2017.04.002

Clark, A. G., Feldman, M. W. & Christiansen, F. (1981). The estimation of epistasis in components of fitness in experimental populations of Drosophila melanogaster I. A two stage maximum likelihood model. Heredity, 46, 321-346.

Borisov, A. Y., Madsen, L. H., Tsyganov, V. E., Umehara, Y., Voroshilova, V. A., Batagov, A. O., Sandal, N. N., Mortensen, A., Schauser, L., Ellis, N., Tikhonovich, I. A. & Stougaard, J. (2003). The Sym35 gene required for root nodule development in Pisum sativum is an ortholog of Nin from Lotus japonicus. Plant Physiology, 131, 1009-1017.

Aagaard, J., Foldager, L., Makki, A., Hansen, V. & Müller-Nielsen, K. (2017). The efficacy of psychoeducation on recurrent depression: A randomised trial with a two-year follow-up. Nordic Journal of Psychiatry, 71(3), 223-229. https://doi.org/10.1080/08039488.2016.1266385

de Silva, E., Thorne, T., Ingram, P., Agrafioti, I., Swire, J., Wiuf, C. & Stumpf, MPH. (2006). The effects of incomplete protein interaction data on structural and evolutionary inferences. BMC Biology, 4(39). https://doi.org/10.1186/1741-7007-4-39

Christiansen, F. (1989). The effect of population subdivision on multiple loci without selection. In M. W. Feldman (Ed.), Mathematical Evolutionary Theory (pp. 71-85). Princeton University Press.

Schierup, M. H., Jensen, T. M. & Wiuf, C. (2005). The effect of a single recombination event. Poster session presented at Genomic Studies and the HapMap, Oxford, United Kingdom.

Bolvig, A. K., Nørskov, N., Hedemann, M. S., Foldager, L., McCarthy-Sinclair, B., Marco, M., Lærke, H. N. & Knudsen, K. E. B. (2017). The effect of antibiotics and diet on enterolactone concentration and metabolome studied by targeted and non-targeted LC-MS metabolomics. Journal of Proteome Research, 16(6), 2135-2150. https://doi.org/10.1021/acs.jproteome.6b00942

Bataillon, T., Mailund, T., Thorlacius, S., Steingrimsson, E., Rafnar, T., Halldorsson, M., Calian, V. & Schierup, M. H. (2006). The effective size of the Icelandic population and the prospects for LD mapping: inference from unphased microsatellite markers. European Journal of Human Genetics, 14, 1044–1053. https://doi.org/10.1038/sj.ejhg.5201669

Benito, B. M., Svenning, J.-C., Kellberg-Nielsen, T., Riede, F., Gil-Romera, G., Mailund, T., Kjaergaard, P. C. & Sandel, B. S. (2017). The ecological niche and distribution of Neanderthals during the Last Interglacial. Journal of Biogeography, 44(1), 51–61. https://doi.org/10.1111/jbi.12845

Bulik, C. M., Thornton, L. M., Parker, R., Kennedy, H., Baker, J. H., MacDermod, C., Guintivano, J., Cleland, L., Miller, A. L., Harper, L., Larsen, J. T., Yilmaz, Z., Grove, J., Sullivan, P. F., Petersen, L. V., Jordan, J., Kennedy, M. A. & Martin, N. G. (2021). The Eating Disorders Genetics Initiative (EDGI): study protocol. BMC Psychiatry, 21(1), Article 234. https://doi.org/10.1186/s12888-021-03212-3

Christiansen, F. (1987). The deviation from linkage equilibrium with multiple loci varying in a stepping-stone cline. Journal of Genetics, 66, 45-67.

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